When there are multiple layers and you display footage in the Player, or play out to tape, or render, Lustre's default behaviour (called Top Vertical Priority) is to display the visible shots from a top-down view of the flattened timeline. This behaviour is further customized by muting layers and prioritizing individual shots. It is also possible to override Top Vertical Priority by soloing a single layer, which forces Lustre to display, play out to tape, and render only that layer.
When selecting segments in the Multi-Layer Timeline, the Storyboard view shows the same selection. See Selecting Shots in the Storyboard.