There are two ways to make tubes react to forces:
Use brush settings to make tubes react to forces
You can make strokes react to forces such as gravity, deflection, and momentum by applying the settings under Tubes > Behavior > Forces. For details, see Set force attributes. Keyframe these settings to make the tubes move dynamically in reaction to the forces.
You can also add turbulence to brushes. Simply turn turbulence on for a stroke, and Paint Effects automatically animates the reaction of the tubes.
To apply turbulence to brushes
Use control curves to make tubes react to forces
If you want stroke tubes to react to forces other than the forces available in the Paint Effects brushes (or in addition to), you can do so by creating control curves and making the control curves react to the forces in your scene. This is an effective technique, for example, for making long hair move naturally or for adding localized forces to trees.
To animate tube behavior by applying forces to control curves